Research Materials of Butterflies in China

This collected works contains 42 papers published in Chinese journals in recent years. It may be used as research materials in the field.

1.
Notes On The Genus Capila From China With A New Species And A New Record (Lepidoptera,Hesperiidae)
中国大弄蝶属一新种及一新纪录种(鳞翅目,弄蝶科)(英文)

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2.
Redescription and Biology of an Endemic Chinese Butterfly: Pieris kozlovi (Alpheraky,1897) (Lepidoptera: Pieridae)
中国特有蝶种——库茨粉蝶(鳞翅目:粉蝶科)再描述及生物学初报(English)
By Li Zhaohui
The endemic Chinese pierid butterfly,Pieris kozlovi(Alpheraky,1897),is redescribed and illustrated in detail based
upon materials collected from the location of holotype in Qinghai autonomous region,northwestern of China.The
grasses Meconopsis spp.of Papaveraceae,Taraxacum spp.of Compositaeand Thalictrum spp.of Ranunculaceae are
recorded as its main nectar source plants.

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3.
Complete mitogenome of the Lesser Purple Emperor Apatura ilia(Lepidoptera:Nymphalidae:Apaturinae) and comparison with other nymphalid butterflies
柳紫闪蛱蝶线粒体基因组全序列及与相关蛱蝶类的比较分析(English)
By Chen Mei
The complete mitochondrial genome of Apatura ilia(GenBank accession no.JF437925) was determined as a circular DNA molecule of 15 242 bp,
with common genes of 13 putative proteins,2 rRNAs,and 22 tRNAs and of the same gene arrangement as in other sequenced lepidopterans.
All protein-coding genes had the typical start codon ATN,except for the COI’s using CGA as its start codon as previously demonstrated in
other lepidopteran species.The comparison of the nucleotide sequences of the A.ilia mitogenome

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4.
A New Species And A New Subspecies Of Lethe HBner(Lepidoptera,Nymphalidae,Satyrinae) From China
中国黛眼蝶属一新种和一新亚种记述(鳞翅目,蛱蝶科,眼蝶亚科)(English)
By Zhai Qing
A new species,Lethe tingeda sp. nov. and a new subspecies,Lethe proxima baoxingensis ssp. nov. from China are described.

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5.
Checklist of the Skipper Genus Pelopidas (Lepidoptera:Hesperiidae) from China with Description of a New Species
中国谷弄蝶属Pelopidas Walker分类并记一新种(鳞翅目:弄蝶科) (English)
By Yuan Xiangqun
The skipper genus Pelopidas contains 11 species worldwide. 7 species are recorded in China. Pelopidas grisemarginata from Ningqiang
County,Shaanxi Province,China is described and illustrated as a new species.

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6.
A Taxonomic Study on the Genus Boeberia Prout,1901 (Lepidoptera:Nymphalidae:Satyrinae) from China
中国贝眼蝶属分类研究(鳞翅目:蛱蝶科:眼蝶亚科) (English)
By Zhai Qing
The genus Boeberia Prout,1901 from China is reviewed. A previously known species is redescribed: Boeberia parmenio(Bber,1809)
and one species is synonymized: Boeberia polyommata Chou and Yuan,2001(=Satyrus ferula(Fabricius,1793)) . Illustrations of adults
and male genitalia are provided.

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7.
A Taxonomic Study on the Genus Boeberia Prout,1901 (Lepidoptera:Nymphalidae:Satyrinae) from China
中国贝眼蝶属分类研究 (鳞翅目:蛱蝶科:眼蝶亚科)*******************************************************************************************************************************************
8.
Identity Of Palaeonympha Avinoffi Schaus,A Satyrine Butterflyfrom China (Lepidoptera,Nymphalidae)
中国古眼蝶的鉴别(鳞翅目,蛱蝶科)
Identity of the Chinese satyrine butterfly Palaeonympha avinoffi Schaus, 1927 has been uncertain since its description.
The holotype is illustrated, and reasons are presented for considering it to represent the same species as Sinonympha amoena
Lee, 1974, new synonym. Sinonympha avinoffi is a new combination. The systematic placement of Sinonympha is yet uncertain.

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9.
Discovery Of The Genus Praescobura Devyatkin (Lepidoptera,Hesperiidae) In China
毗弄蝶属在中国的首次发现(鳞翅目,弄蝶科)(English)
By Fan Xiaoling
Praescobura Devyatkin and P.chrysomaculata are reported from Nanling Nature Reserve in Guangdong and Maoershan Nature Reserve
in Guangxi respectively, as new to China, and female of the species is described for the first time.

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10.
A New Species of the Genus Hyarotis Moore (Lepidoptera: Hesperiidae) from China
中国希弄蝶属Hyarotis Moore一新种(鳞翅目:弄蝶科)(English)
By Fan Xiaoling
A new species of the genus Hyarotis Moore,i.e.,H. quinquepunctatus sp.nov. from Yinggeling of Hainan,China is described and illustrated.
The new species is similar to H. microsticta,but can be easily distinguished from the latter by the two end-cell spots and uncus of male
genitalia with two small processes at distal 1/4 and truncated end.

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11.
Checklist Of The Genus Halpe (Lepidoptera,Hesperiidae) From China With Description Of A New Species
中国酣弄蝶属名录与一新种记述(鳞翅目,弄蝶科)(English)
By Yuanfeng
The species of the skipper genus Halpe Moore occurring in China is brie fly reviewed. The genus Halpe contains 16 known species in China.
Halpe co ncavim arginata is described and illustrated as a new species from Lushan, Sichuan Pr ov ince. The type specimens are deposited
in the Entomological Museum, Northwest Sc i-Tech University of Agriculture and Forestry. A key to species of the genus Halpe from China is
provided for the convenience of identification.

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12.
Notes On The Genus Capila From China With A New Species And A New Record (Lepidoptera,Hesperiidae)
中国大弄蝶属一新种及一新纪录种(鳞翅目,弄蝶科)(English)
By Fanxiao
A new species and a new record of the genus Capila from China are described; i.e. C. pseudomeia sp. nov. from Guangdong and C. phanaeus
Hewitson from Yunnan. A key to the Chinese species of the genus is presented. Type specimens are deposited in the Insect Collection of
Zhongshan University.

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13.
Some New Rhopalocera From Southwest And Northwest China (Lepidoptera: Rhopalocera)
中国西南和西北的一些新蝶类(English)
By Cunshan Xiuyi

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14.
Description Of The Third Species Of Sasakia (Lepidoptera:Nymphalidae)
紫蛱蝶属第三种的记述(鳞翅目:蛱蝶科)(English)
Zhou Yao

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15.
A New Species of the Genus Capila Moore (Lepidoptera: Hesperiidae)
大弄蝶属Capila一新种(鳞翅目:弄蝶科)(English)
By Xiaoling
new species Capila neolineata, sp. n. from Nanling Natural Reserve, Guangdong Province is described and illustrated. This species can be
separated from the related species Capila lineata Chou et Gu, 1994 from Hainan by larger size, broader wing markings and male genitalia.

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16.
A Newly Recorded Genus and Species of the Skippers from China (Lepidoptera: Hesperiidae)
中国弄蝶一新记录属、种(鳞翅目:弄蝶科)(English)
Bu Xiaolin
One genus and species of the skippers are reported as new to China, i.e., Darpa Moore and D. hanria Moore from Mt. Mayuling, Baisha County,
Hainan Province.

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17.
Notes On The Genus Lobocla Moore With Description Of A New Species (Lepidoptera, Hesperiidae)
带弄蝶属研究(鳞翅目,弄蝶科)(English)
By Fan Xiaoling
The species of the genus Lobocla Moore occurring in China is briefly reviewed. The genus contains 8 species. Lobocla quadripunctata is
described and illustrated as a new species from Shennongjia, Hubei Province. Female genitalia of three species are added for the first
time. The type specimens are deposited in the Insect Collection of South China Agricultural University,Guangzhou,China.

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18.
A New Species Of The Genus Deudorix Hewitson From China (Lepidoptera:Rhopalocera)
中国玳灰蝶属一新种(鳞翅目:灰蝶科)(English)
By Wang Min
A new speces of Lycaenidae, Deudorix pseudorapaloides Wang et Chou from Guangki Province is described and illustrated. The new species
is allied to Deudorix rapaliodes Naritomi from Taiwan Province, China, but can be separable from the latter by wing pattern and male genitalia.

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19.
A Revision Of The Genus Gonerilia ShirOZu Et Yamamoto (Lepidoptera: Lycaenidae)
工灰蝶属的修订(鳞翅目:灰蝶科)(English)
onerilia, an endemic genus confined to China, is revised. A new species G. pesthis from Mt. Qinling in Shaanxi Province is described
and illustrated. Five species are included in the genus so far, e.g., G. seraphim, G. thespis, G. buddha, G. okamurai, G. pesthis.
Information on distribution and host plants of the genus is summerized, together with the discussion of its systematic position.

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20.
A Taxonomic Study On The Genus Protacheron Of Ascalaphidae (Neuroptera,Myrmeleontoidea)
蝶角蛉科(脉翅目,蚁蛉总科)中国一新纪录属记述)(English)
By Sun Mingxia
Protacheron Weele is recorded from China for the first time.Protacheron philippinensis(Weele) is recorded for the first time from
China.P.guangxiensis sp.nov.is described from China (Guangxi).The last two species are described and illustrated,and a key to
the three species is given for their separation.

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21.
A Taxonomic Study On The Genus Baes Navs From China(Neuroptera,Psychopsidae)
中国巴蝶蛉属分类研究(脉翅目,蝶蛉科)(English)
By Wang Xinli
The Chinese species of psychopsid genus Balmes Navás, 1910 is revised. Five species are recognized: Balmes birmanus Maclachlan
,B. chikuni sp. nov., B. formosus (Kuwayama), B. notabilis Navás and B. terissinus Navás. A checklist and a key to the Chinese known
species of Balmes are provided. Balmes chikuni sp. nov. is described and illustrated.

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22.
Identity Of Palaeonympha Avinoffi Schaus,A Satyrine Butterflyfrom China (Lepidoptera,Nymphalidae)
中国古眼蝶的鉴别(鳞翅目,蛱蝶科)(English)
Brian Harris
Identity of the Chinese satyrine butterfly Palaeonympha avinoffi Schaus, 1927 has been uncertain since its description. The holotype is
illustrated, and reasons are presented for considering it to represent the same species as Sinonympha amoena Lee, 1974, new synonym.
Sinonympha avinoffi is a new combination. The systematic placement of Sinonympha is yet uncertain.

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23.
Early Stages of Yamamotozephyrus kwangtungensis (Forster) (Insecta,Lepidoptera,Lycaenidae) with Notes on Its Systematic Position
虎灰蝶幼期及分类地位)(English)
By Wangmin
The genus Yamamotozephyrus Saigusa is a monobasic genus with Y.kwangtungensis(Forster) as the type species.Early stages of the
genus are reported for the first time from the Nanling Mountains,S.China.The food plant of the species is Castanopsis sp.The species
has one generation per year,with adults flying in May and June.

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24.
New Species, Subspecies and New Records of Butterflies (Lepidoptera) from China, I
中国蝴蝶新种、新亚种及新记录种(I)(鳞翅目))(Chinese with English Summary)
By Zhou Yao
In the present paper, one new species and two new subspecies with new records of two species and two subspecies are described from
Sichuan, Yunnan and Hainan Provinces of China. All the specimens are deposited in the Entomological Museum, Northwest Sci Tech
University of Agriculture and Forestry. Papilionidae 1. Papilio pavonis, sp. n. (Figs.1～2) This species is similar to P.
polyctor Boisduval, butdistinguishable from the latter in the following aspects: a)forewing without green cross band

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25.
New Species, Subspecies and New Records of Butterflies (Lepidoptera) from China, II
中国蝴蝶新种、新亚种及新记录(II)(鳞翅目:凤蝶科) (Chinese with English Summary)
By Zhou Yao

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26.
New Species New subspecies and New Record of Butterflies (Lepidoptera) from China, Ⅲ
中国蝴蝶新种、新亚种及新记录种(Ⅲ)(鳞翅目)(Chinese with English Summary)
By Zhou Yao
In the present paper, 1 new species, 4 new subspecies, 1 new form and 1 new record species of Pieridae from China, and reports 6
known species,these were not described in 《Monographia Rhopalocerorum Sinensium》. All type specimens are kept in the Entomological
Museum Northwestern Sci Tech University of Agriculture and Forestry. Pieridae 1. Colias aurora Esper f. chloe Eversman, (Figs.1,2)
Obs. specimen: 1♀, Pingliang, Gansu, 15 Ⅶ 1992, col. Wang Huanran. Not described in 《Monogr. Rhop. Sin.》.

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27.
New Species, New Subspecies and New Record of Butterflies (Lepidoptera) from China (IV)
中国蝴蝶新种新亚种及新记录种(IV)(鳞翅目)(Chinese with English Summary)
By Zhou Yao
In the present paper 3 new species, 3 new subspecies and l american invasive species are described from Hainan, Fujian, Sichuan,
Qinghai and Shaanxi, China. All the specimens are deposited in the Entomological Museum, Northwest Sci Tech. University of
Agriculture and Forestry. Danaidae 1. Parantica hainanica, sp. nov. (Figs.1,2) This species is simllar to P. aglea (Stoll),
but with only one series of white spots in the lack marginal band of both wings. Holotype: ♀, Lingshui xian, Hainan Provinc...

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28.
New Species,New Subspecies and New Records of Butterflies(Lepidoptera:Satyridae)from China(V)
中国蝴蝶新种,新亚种及新记录种(V)(鳞翅目:眼蝶科)(Chinese with English Summary)
By Zhou Yao
The members of the family Satyridae are usually medium sized or small sized butterflies. The wings are usually dull brown.
They usually are not attracted by collectors.. Wu Chenfu (1938) listed 224 species of Satyridae in China in "Catalogus
Insectorum Sinensium". Li Chuanlong (1958) counted up 246 species n China. Chou Io et al (1994) recorded 212 species in
"Monographia Rhopalocerorum Sinensium".Huang Renxin(2000) added 32 known pecies in "Butterfly of Xinjiang".

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29.
New Species, New Subspecies, and New Records of Butterflies from China (VI)
中国蝴蝶新种、新亚种及新记录种(VI)(Chinese with English Summary)
By Zhou Yao
The present paper deals with descriptions of 12 new species, 2 new subspecies, 1 new form, 6 new record species of Nymphalidae from China
and reports 4 known species, which were not included in"Monographia Rhopalocerorum Sinensium". All the type specimens are kept in the
Entomological Museum, Northwest Sci-Tech University of Agriculture and Forestry. 1. Cethosia rubra, sp. nov. (Figs. 1a, 1b) This species is
similar to C. biblis Drury, but fore wings with a white outer medianline maculae at ape

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30.
Descriptions of two New Subspecies of the Genus Antigius Sibatani et Ito(Lepidoptera: Lycaenidae) from China
中国青灰蝶属Antigius Sibatani et Ito二新亚种描述( English )
By Wang Min
Two new subspecies of the genus Antigius Sibatani et Ito are described from Zhengjiang and Guangdong Provinces.A.cheni zhengjiangensis
ssp.n.is separated from the nominate subspecies from Sichuan by the less developed white marking on hind wing and the broader distal cell
streaks on wing underside.A.butleri nanlingensis ssp.n.is distinguished from the nominate subspecies by the less developed white marking
on hindwing upperside,much broader postdiscal bands on undersides of both wings and the di

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31.
Studies on the butterfly fauna in Luo Mountain National Nature Reserve of Ningxia,China
宁夏罗山国家自然保护区蝶类区系研究(Chinese with English Summary)
By Ji Xuelong
The butterfly fauna in Luo Mountain Nature Reserve were investigated during from 2008 to 2009.The results show that there are 45 species
butterflies belonging to 6 families and 32 genera in this area.The family Nymphalidae is the predominant group(8 genera and 14 species),
occupying 28.13 % of the total species.The family Lycaenidae(9 genera and 10 species)and Satyridae(8 genera and 9 species) is the next,
accounting for 25%.respectively,of the total.Analysis on the faunal composition from

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32.
The diversity and zonal distribution of butterflies in Shanxi Province
山西省蝶类多样性与地带分布(Chinese with English Summary)
By Cao Tianwen
On the basis of natural geography and vegetation, the present situation of butterfly diversity was studied in Shanxi. The results indicated
that:in different forest regions, the diversity of butterfly had much difference and decreased progressively from south to north. Meanwhile,
influenced by the microclimate environment, the butterfly resources appeared clumpy distribution. In cultivation area, the diversity of
species lost seriously. Based on the distribution of 216 species of butterflies in...

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33.
Composition And Fauna Analysis Of Butterflies In Western Gansu Province
甘肃西部蝶类区系研究(Chinese with English Summary)
By Guan Puguo
More than 1200 specimens of butterflies, which belong to 8 families, 73 genera, 127 species and subspecies, were collected in the
western ansu from 1994 to 1998. Among them, 44 species and subspecies are new recorded in Gansu province. A total of 74 species
of which accounted to 58.8% belong to the palaearctic realm, the other 53 species making up 41.2% are widely scattered in the
oriental realm and alaearctic realm.

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34.
The New Record of Butterflies in Gansu Province
甘肃省蝶类新记录(Chinese with English Summary)
By Tang Chunmei
The paper recorded 25 species of butterflies collected in Tianshui Area of Gansu;they belong to 20 genera of 3 families.Among them 2 species
of Nymphalidae,17 species of Lycaenidae,6 specie of Hesperiidae.

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35.
Study on fauna and utilization of butterflies in Hunan Province
湖南蝶类区系及利用研究(Chinese with English Summary)
By Zhu Xun
There are 158 species of butterflies in Hunan Province,the elements of fauna are major Oriental,secondary Paleo-artic one.In order to develop
economy of Hunan,this paper gives 4 aspects of suggestion on how to strenghthen the study of exploitation and utilization of butterflies.

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36.
Investigation on Butterfly in Southwest Mountain Area of Zhejiang
浙西南山区蝴蝶种类调查(Chinese with English Summary)
By Zhao Renyou
Investigations on butterflies in southwest mountain area in Zhejiang demonstrated that there has 169 species of 10 families, among which 19
Papilionidae, 3 Amathusiidae, 12 Pieridae, 30 Satyridae,54 Nymphalidae, 25 Hesperiidae, 22 Lycaenidae, 1 Acraeidae, 2 Riodinidae, 1
Danaidae. There are five new records in Zhejiang and one of them needs to be named, and four types are under the state protection.

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37.
Investigation on Butterfly Faunas in Baikal and Jiangsu Regions
贝加尔与江苏蝶类区系比较及成因分析(Chinese with English Summary)
By Cui Miao
The investigations of the butterfly fauna were carried,and near 200 specimens and more than 1000 specimens were collected respectively in
Baikal and Jiangsu regions.The butterfly faunas of the two regions are provided in the present paper on the basis of the investigations in the
two regions combined with the works of previous researchers,including 178 species belonging to 10 families and 105 genera in Jiangsu and
55 species belonging to 7 families and 39 genera in Baikal.

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38.
The Taxa Of The Pieris Napi- Bryoniae Group (Sensu Lato )In China (Lepidoptera: Pieridae)
中国菜粉蝶群(Pieris napi-broniae)的分类(鳞翅目:粉蝶科)(English)
Hou Taoqian
Summary The species and subspecies of the Pieris napi-bryoniae group are listed and where necessary discussed in greater detail.

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39.
Comparison of Nucleic Acid Content among Non-diapause Pupae, Diapause Pupae and Eclosion-adult from Diapause Pupae of Papilio memnon
美凤蝶非滞育蛹·滞育蛹与滞育蛹羽化成虫核酸含量比较(English)
By Yi Chuanhui
[Objective] The aim of this study was to provide basis for deeply understanding the diapause mechanism of Papilio memnon L. [Method] RNA and DNA
content of non-diapause pupae, diapause pupae and eclosion-adult from diapause pupae at different development stages were detected by the
colorimetry. [Result] RNA content of non-diapause pupae was 4.614 0-7.946 3 μg/mg, while diapause pupae was 4.326 0-5.885 3 μg/mg and
eclosion-adult from diapause pupae was 20.779 3 μg/mg at initial stage. DNA content

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40.
Comparison of Protein Content among Non-diapause Pupae,Diapause Pupae and Eclosion-adult from Diapause Pupae of Papilio memnon
美凤蝶非滞育蛹与滞育蛹及其羽化成虫蛋白质含量比较(English)
By Yi Chuanhui
[Objective]The experiment aimed to explore the diapause mechanism of Papilio memnon L.further.[Method] The colorimetry method was used to
detect protein content of non-diapause pupae,diapause pupae and eclosion-adult from diapause pupae at different developmental stages.[Result]
It was showed that the protein content of non-diapause pupae was from 111.768 to 115.030 7 mg/g and the change of protein content during whole
pupal stage,while diapause pupae were from 81.218 to 131.623 7 mg/g and the c

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41.
Phylogeny of Princeps butterflies from China: evidences from mitochondrial partial COⅠ and COⅡ gene sequences (Lepidoptera: Papilionidae)
从COⅠ和COⅡ基因部分序列研究中国翠凤蝶亚属(鳞翅目:凤蝶科)的分子系统关系(English)
In order to study phylogenetic relationships among Princeps species of China, partial COⅠ (about 627 bp) and COⅡ (about 690 bp) gene sequence from 10
species of Princeps in China were analyzed. There were 184 variable sites and 144 parsimony-informative sites in the 1 317 nucleotides of the data,
Princeps exhibited a 74.2 Arginine and Thymine bias. Molecular phylogenetic trees were reconstructed using maximum parsimony (MP) method
and maximum likelihood (ML) method. The molecular phylogenetic

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42.
Phylogenetic evaluation of the taxonomic status of Papilio maackii and P.syfanius(Lepidoptera:Papilionidae)
基于系统发生探讨绿带翠凤蝶和西番翠凤蝶的分类地位(English)
By Chu Lixin
The taxonomic status of Papilio maackii and P.syfanius has long been disputed.We conducted a molecular phylogenetic study to evaluate the
taxonomic status of P.maackii and P.syfanius.A total of twenty-four P.maackii individuals from six localities and sixteen P.syfanius individuals
from two localities were analyzed.We sequenced the partial region of the CO-I gene(about 579 bp) and partial CO-II gene sequence(about 655bp) of the two species.
The Kimura-2-Parameter distances among P.maackii

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